# positions[].organizations[].parent[].label
College of Veterinary Medicine and Biomedical Sciences

# positions[].organizations[].parent[].label
School of Veterinary Medicine and Biomedical Sciences

# overview
Dr. Dickson Varner is a Diplomate of the American College of Theriogenologists. His special interests include equine theriogenology, fertility probes for stallions, in-vitro preservation of equine spermatozoa, capacitation of equine spermatozoa, assisted reproductive techniques, and subfertility in stallions.

# keywords[]
Reverse Transcriptase Polymerase Chain Reaction

# keywords[]
30 Agricultural, Veterinary And Food Sciences

# keywords[]
1114 Paediatrics And Reproductive Medicine

# keywords[]
Insulin-like Growth Factor Binding Protein 5

# keywords[]
Insulin-like Growth Factor Binding Protein 2

# keywords[]
4606 Distributed Computing And Systems Software

# keywords[]
30 Agricultural, Veterinary And Food Sciences

# keywords[]
30 Agricultural, Veterinary And Food Sciences

# keywords[]
Clinical Trials And Supportive Activities

# keywords[]
30 Agricultural, Veterinary And Food Sciences

# keywords[]
30 Agricultural, Veterinary And Food Sciences

# keywords[]
30 Agricultural, Veterinary And Food Sciences

# keywords[]
Cyclic Nucleotide Phosphodiesterases, Type 3

# keywords[]
1114 Paediatrics And Reproductive Medicine

# keywords[]
Reverse Transcriptase Polymerase Chain Reaction

# educationAndTraining[].organization.label
Texas A&M University - (College Station, Texas, United States)

# educationAndTraining[].organization.label
University of Pennsylvania - (Philadelphia, Pennsylvania, United States)

# educationAndTraining[].organization.label
University of Missouri - (Columbia, Missouri, United States)

# educationAndTraining[].organization.label
University of Missouri - (Columbia, Missouri, United States)

# publications[].label
Calcium-calmodulin and pH regulate protein tyrosine phosphorylation in stallion sperm.

# publications[].label
Effect of holding technique and culture drop size in individual or group culture on blastocyst development after ICSI of equine oocytes with low meiotic competence.

# publications[].label
Effect of calcium, bicarbonate, and albumin on capacitation-related events in equine sperm.

# publications[].label
Efficiency of spermatogenesis: a comparative approach.

# publications[].label
In vitro-produced equine embryos: production of foals after transfer, assessment by differential staining and effect of medium calcium concentrations during culture.

# publications[].label
Apoptosis during the initiation of equine spermatogenesis

# publications[].label
Chromatin configuration within the germinal vesicle of horse oocytes: changes post mortem and relationship to meiotic and developmental competence.

# publications[].label
Advances in cooled semen technologies: seminal plasma and semen extender.

# publications[].label
The effect of two levels of hemospermia on stallion fertility.

# publications[].label
Fertilizing capacity of equine spermatozoa stored for 24 hours at 5 or 20 degrees C.

# publications[].label
Use of a computerized system for evaluation of equine spermatozoal motility.

# publications[].label
Effect of co-culture with theca interna on nuclear maturation of horse oocytes with low meiotic competence, and subsequent fusion and activation rates after nuclear transfer.

# publications[].label
Use of a silane-coated silica particle solution to enhance the quality of ejaculated semen in stallions

# publications[].label
The impact of cushioned centrifugation protocols on semen quality of stallions.

# publications[].label
Sperm DNA quality evaluated by comet assay and sperm chromatin structure assay in stallions after unilateral orchiectomy.

# publications[].label
Evaluation and diagnosis of acrosome function/dysfunction in the stallion.

# publications[].label
Relationship between sperm nuclear protamine free -SH status and susceptibility to DNA denaturation.

# publications[].label
Agreement between measures of total motility and membrane integrity in stallion sperm.

# publications[].label
Horse Y chromosome assembly displays unique evolutionary features and putative stallion fertility genes.

# publications[].label
Validation of a commercially available fluorescence-based instrument to evaluate stallion spermatozoal concentration

# publications[].label
The effect of flash-freezing temperature on stallion sperm DNA structure.

# publications[].label
Measurements of glycosaminoglycans in follicular, oviductal and uterine fluids of mares.

# publications[].label
Granulocyte-macrophage colony-stimulating factor reduces testicular germ cell apoptosis in vitro

# publications[].label
Azoospermia in stallions: determining the cause.

# publications[].label
Mitochondrial oxygen consumption is a unique indicator of stallion spermatozoal health and varies with cryopreservation media.

# publications[].label
Genome-wide association study implicates testis-sperm specific FKBP6 as a susceptibility locus for impaired acrosome reaction in stallions.

# publications[].label
Bilateral infectious epididymitis in a stallion.

# publications[].label
Relationship of seminal plasma level and extender type to sperm motility and DNA integrity.

# publications[].label
Induction and characterization of acrosome reaction in equine spermatozoa.

# publications[].label
Semen Processing for the Subfertile Stallion

# publications[].label
Case Study of Processing and Insemination Techniques: Attempts to Improve Fertility of an Aged Stallion with Dilute Semen of Poor Quality

# publications[].label
Theriogenology question of the month. Scrotal hematocele.

# publications[].label
The relationship of daily sperm production with number of Sertoli cells and testicular size in adult horses: role of primitive spermatogonia.

# publications[].label
Effect of non-sperm cells removal with single-layer colloidal centrifugation on myeloperoxidase concentration in post-thaw equine semen.

# publications[].label
Induction of the acrosome reaction in stallion spermatozoa: effects of incubation temperature, incubation time, and ionophore concentration

# publications[].label
Focal adhesion kinases and calcium/calmodulin-dependent protein kinases regulate protein tyrosine phosphorylation in stallion sperm.

# publications[].label
Stallion karyotype variability and lack of association with abortion: a case report.

# publications[].label
Effects of cryopreservation on the acrosomal status of stallion spermatozoa.

# publications[].label
POSTPARTURIENT HEMORRHAGE IN THE MARE - MANAGING RUPTURED ARTERIES OF THE BROAD LIGAMENT

# publications[].label
Stallion sperm transcriptome comprises functionally coherent coding and regulatory RNAs as revealed by microarray analysis and RNA-seq.

# publications[].label
Effect of ovary storage and oocyte transport method on maturation rate of horse oocytes.

# publications[].label
Factors affecting developmental competence of equine oocytes after intracytoplasmic sperm injection.

# publications[].label
Pregnancy rates in mares following hysteroscopic or transrectally-guided insemination with low sperm numbers at the utero-tubal papilla.

# publications[].label
Regulation of axonemal motility in demembranated equine sperm.

# publications[].label
A Bayesian approach to prediction of stallion daily sperm output.

# publications[].label
Effects of bilateral ovariectomy via colpotomy in mares: 23 cases (1984-1990).

# publications[].label
Effect of surgical technique for unilateral orchiectomy on subsequent testicular function in Miniature Horse stallions.

# publications[].label
Production of live foals via intracytoplasmic injection of lyophilized sperm and sperm extract in the horse.

# publications[].label
Breeding with cooled, transported equine semen: The insemination process

# publications[].label
Relationship of Sperm Quality to Fertility after 4 Days of Cooled Storage of Equine Semen

# publications[].label
Effect of daily semen centrifugation and resuspension on the longevity of equine sperm quality following cooled storage.

# publications[].label
Elective cesarean section in mares: eight cases (1980-1989).

# publications[].label
Stabilization of extragonadal sperm reserves in stallions: application for determination of daily sperm output

# publications[].label
Sperm DNA assays and their relationship to sperm motility and morphology in bulls (Bos Taurus).

# publications[].label
Dexamethasone downregulates expression of several genes encoding orphan nuclear receptors that are important to steroidogenesis in stallion testes.

# publications[].label
Presumptive evidence of a preovulatory sperm reservoir in the mare: morphological investigations using scanning electron microscopy

# publications[].label
Lactate and Pyruvate Are Major Sources of Energy for Stallion Sperm with Dose Effects on Mitochondrial Function, Motility, and ROS Production.

# publications[].label
Use of Piezo-driven direct nuclear injection and activation with stallion sperm extract to produce horse nuclear transfer embryos

# publications[].label
Fertility comparison between breeding at 24 hours or at 24 and 48 hours after collection with cooled equine semen.

# publications[].label
Evaluating breeding soundness in stallions .2. Semen collection and evaluation

# publications[].label
The addition of ticarcillin-clavulanic acid to INRA 96 extender for stallion semen cooling.

# publications[].label
The "dilution effect" in stallion sperm.

# publications[].label
Characterization of Equine Phospholipase C Zeta: A Review and Preliminary Results on Expression Defects in Subfertile Stallions

# publications[].label
Effect of extender pH on motility characteristics of cool-stored equine spermatozoa

# publications[].label
Equine sperm and oviductal epithelia (uterine tubal) interactions in co- culture

# publications[].label
Blastocyst development in equine oocytes with low meiotic competence after suppression of meiosis with roscovitine prior to in vitro maturation.

# publications[].label
Effects of holding time and media on meiotic and developmental competence of horse oocytes

# publications[].label
In vitro fertilization of in vitro-matured equine oocytes: effect of maturation medium, duration of maturation, and sperm calcium ionophore treatment, and comparison with rates of fertilization in vivo after oviductal transfer.

# publications[].label
Thermoregulation of the testicle in response to exercise and subsequent effects on semen characteristics of stallions.

# publications[].publisher
American Veterinary Medical Association (AVMA)

# publications[].label
Evaluating breeding soundness in stallions .5. Predicting potential fertility

# publications[].label
Spatio-Temporal Variation in Viability of Spermatozoa of Honey Bee, Apis mellifera, Drones in Central Texas Apiaries

# publications[].label
Diagnostic aids for the detection of urine in the equine ejaculate.

# publications[].label
Approaches to Breeding Soundness Examination and Interpretation of Results

# publications[].label
Apoptosis as a mechanism of germ cell loss in mature stallions

# publications[].label
Intra- and inter-stallion variation in sperm morphology and their relationship with fertility.

# publications[].label
Breeding Management of the Older Stallion With Declining Testicular Function

# publications[].label
Biodegradable estradiol microspheres do not affect uterine involution or characteristics of postpartum estrus in mares.

# publications[].label
Signalment, clinical features, and outcome for male horses with urethral rents following perineal urethrotomy or corpus spongiotomy: 33 cases (1989-2013).

# publications[].publisher
American Veterinary Medical Association (AVMA)

# publications[].label
Breeding with cooled, transported equine semen: Testing and preparing semen for travel

# publications[].label
Effects of postparturient uterine lavage on uterine involution in the mare.

# publications[].label
Breeding with cooled transported equine semen: The advantages and success rate

# publications[].label
Capacitation and acrosome reaction of equine spermatozoa by heparin

# publications[].label
Hysteroscopic or rectally guided, deep-uterine insemination of mares with spermatozoa stored 18 h at either 5 or 15 degrees C prior to flow-cytometric sorting

# publications[].label
Testicular and hormonal changes in stallions with thermally induced testicular degeneration.

# publications[].label
Stallion fertility: a focus on the spermatozoon.

# publications[].label
Effect of intrauterine treatment with prostaglandin E2 prior to insemination of mares in the uterine horn or body.

# publications[].label
Effect of artificial vagina lubricants on stallion sperm quality.

# publications[].label
Use of a semen extender containing antibiotic to improve the fertility of a stallion with seminal vesiculitis due to Pseudomonas aeruginosa.

# publications[].label
Influence of day of postpartum breeding on pregnancy rate, pregnancy loss rate, and foaling rate in Thoroughbred mares.

# publications[].label
EFFECT OF HEPARIN ON CAPACITATION/ACROSOME REACTION OF EQUINE SPERM

# publications[].label
In vitro culture of precision-cut testicular tissue as a novel tool for the study of responses to LH.

# publications[].label
Equine blastocyst development after intracytoplasmic injection of sperm subjected to two freeze-thaw cycles.

# publications[].label
Supplemental Antibiotics in a Commercial Extender for Stallion Semen.

# publications[].label
Dexamethasone acutely regulates endocrine parameters in stallions and subsequently affects gene expression in testicular germ cells.

# publications[].label
Surgical Correction of Abnormalities Affecting the Reproductive Organs of Stallions

# publications[].label
Cholesterol-to-phospholipid ratio in whole sperm and seminal plasma from fertile stallions and stallions with unexplained subfertility.

# publications[].label
Management Options for the Aged Breeding Stallion with Declining Testicular Function

# publications[].label
Use of a modified Vinsot technique for partial phallectomy in 11 standing horses.

# publications[].publisher
American Veterinary Medical Association (AVMA)

# publications[].label
Blastocyst formation rates in vivo and in vitro of in vitro-matured equine oocytes fertilized by intracytoplasmic sperm injection.

# publications[].label
Prepackaging of equine semen in goblets prior to cryopreservation.

# publications[].label
Total RNA isolation from stallion sperm and testis biopsies.

# publications[].label
AN UPDATE ON UTERINE DEFENSE-MECHANISMS IN THE MARE

# publications[].label
Urethral defects in geldings with hematuria and stallions with hemospermia.

# publications[].label
Administration of oxytocin immediately after insemination does not improve pregnancy rates in mares bred by fertile or subfertile stallions.

# publications[].label
Production of cloned horse foals using roscovitine-treated donor cells and activation with sperm extract and/or ionomycin.

# publications[].label
The effect of a surgically created shunt between the corpus cavernosum penis and corpus spongiosum penis of stallions on erectile and ejaculatory function.

# publications[].label
Effects of semen fractionation and dilution ratio on equine spermatozoal motility parameters.

# publications[].label
Techniques for evaluating selected reproductive disorders of stallions.

# publications[].label
Advances in large-animal reproduction: The expanding role of ultrasonography

# publications[].label
Effect of holding at room temperature on initial chromatin configuration and in vitro maturation rate of equine oocytes.

# publications[].label
Effect of cryopreservation protocol on postthaw characteristics of stallion sperm.

# publications[].label
The effect of dual-hemisphere breeding on stallion fertility.

# publications[].label
Effects of seminal plasma and flash-freezing on DNA structure of stallion epididymal sperm exposed to different potentiators of DNA damage.

# publications[].label
Season but not age affects Sertoli cell number in adult stallions.

# publications[].label
Activation of cumulus-free equine oocytes: effect of maturation medium, calcium ionophore concentration and duration of cycloheximide exposure.

# publications[].label
Relative alterations in core body temperature and internal and external scrotal temperatures of exercising stallions

# publications[].label
Insulin-like growth factor-I and insulin-like growth factor binding protein-2 and -5 in equine seminal plasma: association with sperm characteristics and fertility.

# publications[].label
Developments in stallion semen evaluation.

# publications[].label
Evaluating breeding soundness in stallions .1. The basic examination

# publications[].label
Pregnancy Rates in Tranquilized Maiden Thoroughbred Mares

# publications[].label
Use of an ecraseur for ovariohysterectomy in mares.

# publications[].label
Low daily sperm output per ml of testis as a diagnostic criteria for testicular degeneration in stallions

# publications[].label
Efficacy of Long-Acting Formulations of Estradiol or Progesterone Plus Estradiol on Estrous Synchronization in Broodmares

# publications[].label
CatSper and the relationship of hyperactivated motility to intracellular calcium and pH kinetics in equine sperm.

# publications[].label
Advances in large-animal reproduction: Updated techniques for artificial insemination and embryo transfer

# publications[].label
Effect of insemination timing on the fertilizing capacity of frozen/thawed equine spermatozoa.

# publications[].label
Surgical repair of the lacerated cervix in the mare.

# publications[].label
Azoospermia in stallions: determining the cause.

# publications[].label
Priapism in a stallion with generalized malignant melanoma.

# publications[].label
Artificial insemination and preservation of semen.

# publications[].label
Regulation of estrus and ovulation in mares with progesterone or progesterone and estradiol biodegradable microspheres with or without PGF2alpha.

# publications[].label
Cervical wedge resection for treatment of pyometra secondary to transluminal cervical adhesions in six mares.

# publications[].publisher
American Veterinary Medical Association (AVMA)

# publications[].label
Role of Sertoli cell number and function on regulation of spermatogenesis.

# publications[].label
Production of nuclear transfer horse embryos by Piezo-driven injection of somatic cell nuclei and activation with stallion sperm cytosolic extract.

# publications[].label
Hysteroscopic or rectally guided, deep-uterine insemination of mares with spermatozoa stored 18 h at either 5 degrees C or 15 degrees C prior to flow-cytometric sorting.

# publications[].label
Effect of daily spermatozoan production but not age on transit time of spermatozoa through the human epididymis.

# publications[].label
Cholesterol to phospholipid ratio in sperm of stallions with unexplained subfertility.

# publications[].label
Holding immature equine oocytes in the absence of meiotic inhibitors: effect on germinal vesicle chromatin and blastocyst development after intracytoplasmic sperm injection.

# publications[].label
Effect of centrifugal fractionation protocols on quality and recovery rate of equine sperm.

# publications[].label
Comparison of three containers used for the transport of cooled stallion semen.

# publications[].label
Development of Integrated DAO Pattern Applying Iterator Pattern

# publications[].label
Gene expression in the spermatogenically inactive "dark" and maturing "light" testicular tissues of the prepubertal colt.

# publications[].label
The effects of antibiotic type and extender storage method on sperm quality and antibacterial effectiveness in fresh and cooled-stored stallion semen.

# publications[].label
The relationship between mitochondrial DNA copy number and stallion sperm function.

# publications[].label
Dense spermatozoa in stallion ejaculates contain lower concentrations of mRNAs encoding the sperm specific calcium channel 1, ornithine decarboxylase antizyme 3, aromatase, and estrogen receptor alpha than less dense spermatozoa.

# publications[].label
Abortion of a mummified fetus associated with short uterine body in a mare.

# publications[].label
Suppression of meiosis by inhibitors of m-phase proteins in horse oocytes with low meiotic competence.

# publications[].label
USE OF A RAPID PROGESTERONE ASSAY IN A BEEF-CATTLE ESTRUS SYNCHRONIZATION PROGRAM

# publications[].label
Three methods of oxytocin-induced parturition and their effects of foals.

# publications[].label
In vitro interaction between oviduct epithelial and equine sperm.

# publications[].label
Luteinizing hormone and progesterone concentrations and induction of estrus after use of norgestomet ear implants or constant infusion of gonadotropin-releasing hormone in anestrous, nonlactating dairy goats.

# publications[].label
Evaluation of cellulose acetate/nitrate filters for the study of stallion sperm motility.

# publications[].label
Factors impacting equine sperm recovery rate and quality following cushioned centrifugation.

# publications[].label
The effect of postbreeding uterine lavage on pregnancy rate in mares.

# publications[].label
Relationship of seminal plasma level and extender type to sperm motility and DNA integrity

# publications[].label
Strategies for Processing Semen from Subfertile Stallions for Cooled Transport.

# publications[].label
Advances in large-animal reproduction: In vitro production of embryos

# publications[].label
PULSATILE RELEASE OF GONADOTROPINS IN YOUNG PONY STALLIONS

# publications[].label
Penile paralysis and paraphimosis associated with reserpine administration in a stallion.

# publications[].label
Mares with delayed uterine clearance have an intrinsic defect in myometrial function.

# publications[].label
Bilateral seminal vesiculitis and ampullitis in a stallion.

# publications[].label
Effect of age and season on the establishment of spermatogenesis in the horse.

# publications[].label
Pregnancy rates in mares inseminated with 0.5 or 1 million sperm using hysteroscopic or transrectally guided deep-horn insemination techniques.

# publications[].label
Ejaculate and type of freezing extender affect rates of fertilization of horse oocytes in vitro.

# publications[].label
Relationship between stallion sperm motility and viability as detected by two fluorescence staining techniques using flow cytometry.

# publications[].label
Evaluating breeding soundness in stallions .4. Hormonal assay and testicular biopsy

# publications[].label
Bacteriology of preserved stallion semen and antibiotics in semen extenders.

# publications[].label
The effects of urine concentration, and cushion centrifugation to remove urine, on the quality of cool-stored stallion sperm.

# publications[].label
Effect of Antibiotic-containing Extenders on Taylorella equigenitalis Contaminated Semen

# publications[].label
Physiological responses of mature quarter horses to reining training when fed conventional and fat-supplemented diets

# publications[].label
Surgical Correction of Abnormalities Affecting the Reproductive Organs of Stallions

# publications[].label
Effect of feeding a DHA-enriched nutriceutical on the quality of fresh, cooled and frozen stallion semen.

# publications[].label
THE EFFECT OF UTERINE LAVAGE PERFORMED 4 HOURS POST INSEMINATION ON PREGNANCY RATE IN MARES

# publications[].label
Evaluating breeding soundness in stallions .3. Useful laboratory tests

# publications[].label
Effect of storage time and temperature on stallion sperm DNA and fertility.

# publications[].label
Comparison of methods for assessing integrity of equine sperm membranes.

# publications[].label
The effects of growth factor on testicular germ cell apoptosis in the stallion

# publications[].label
Production of horse foals via direct injection of roscovitine-treated donor cells and activation by injection of sperm extract.

# publications[].label
Physiologic responses of acclimatized or non-acclimatized mature reining horses to heat stress: I. Heart rate, respiration rate, lactate, rectal temperature, cortisol and packed cell volume

# publications[].label
Effects of dead spermatozoa on motion characteristics and membrane integrity of live spermatozoa in fresh and cooled-stored equine semen.

# publications[].label
Effect of heparin on capacitation/acrosome reaction of equine sperm.

# publications[].label
Factors affecting spermatogenesis in the stallion.

# publications[].label
The relationship between sperm quality in cool-shipped semen and embryo recovery rate in horses.

# publications[].label
Clinical observations on changes in concentrations of hormones in plasma of two stallions with thermally-induced testicular degeneration

# publications[].label
DNA microarray analysis of gene expression in testicular tissue of stallions

# publications[].label
Effect of cholesterol on the motility and plasma membrane integrity of frozen equine spermatozoa after thawing.

# publications[].label
Effects of cooling rate and storage temperature on equine spermatozoal motility parameters.

# publications[].label
Developmental competence in vivo and in vitro of in vitro-matured equine oocytes fertilized by intracytoplasmic sperm injection with fresh or frozen-thawed spermatozoa.

# publications[].label
Effects of gas conditions, time of medium change, and ratio of medium to embryo on in vitro development of horse oocytes fertilized by intracytoplasmic sperm injection.

# publications[].label
Germ cell apoptosis in the testes of normal stallions.

# publications[].label
Use of prostaglandin E2 to ripen the cervix of the mare prior to induction of parturition.

# publications[].label
Effects of ecbolic agents on measurements of uterine involution in the mare.

# publications[].label
Some Factors Associated With Fertility of Thoroughbred Stallions

# publications[].label
A comparison between cervical dimensions of pregnant and nonpregnant Santa Gertrudis and Bos taurus cows.

# publications[].label
Breeding with cooled, transported equine semen: Proper mare management

# publications[].label
Dexamethasone acutely down-regulates genes involved in steroidogenesis in stallion testes.

# publications[].label
Recommendations for clinical GnRH challenge testing of stallions

# publications[].label
CELL-TO-CELL COMMUNICATION OF EQUINE UTERINE TUBE (OVIDUCT) CELLS AS DETERMINED BY ANCHORED CELL ANALYSIS IN CULTURE

# publications[].label
Effects of transport container and ambient storage temperature on motion characteristics of equine spermatozoa.

# streetAddress
Texas A&M University; Large Animal Clinical Sciences; 4475 TAMU

# organizations[]
College of Veterinary Medicine and Biomedical Sciences

# organizations[]
School of Veterinary Medicine and Biomedical Sciences

# schools[]
College of Veterinary Medicine and Biomedical Sciences

# schools[]
School of Veterinary Medicine and Biomedical Sciences

# inTheNews[].label
Dr. Dickson Varner Joins Texas Tech School as Visiting Professor